From PBS Eons.

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How did the "false horse," Thoatherium, and its relatives survive when their hoofed legs seemed to be adapted for an ecosystem that wouldn’t exist for another 12 million years?

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References:
https://docs.google.com/document/d/1NR3OQfiULdz7Ohblh6a0vRyopP6QxqD8q7dNzbz8Ic4/edit?usp=sharing