From NPR.

A beloved Galápagos tortoise named Gramma, the oldest resident of the San Diego Zoo, has died at an estimated age of 141 years old.

Gramma first came to San Diego from the Bronx Zoo nearly a century ago and quickly became known for her gentle, shy personality. Keepers affectionately called her “the queen of the zoo,” and visitors adored her for decades.

While Galápagos tortoises can live well over 100 years — and even longer under human care — Gramma had been struggling with age-related bone issues before she was euthanized.